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I ink the whole page, putting in outlines only. I've added black only where I know I'll need to ink in grey - otherwise it would be hard to see what I was doing.
It is possible to ink in black, put the shadows underneath and convert the outlines over the shadows to grey, but it can be fiddly. It's also possible to draw in black and set the blending mode of the outlines (or layer in Photoshop and Painter) to Exclusion (Reverse Out in Painter), but the effect can be unpredictable.